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Table 2 The following table gives a summary of all long-term studies we found that quantified changing fuel loads and/or forest cover over time. Some of these studies are not included in Fig. 1 if there were other studies in the same location that provided the same information or covered a longer period of time

From: Trends in western USA fire fuels using historical data and modeling

Title

Author and year published

Region

Veg type

Years of study

Disturbance during study

Summary

URL

Landscape Changes in the Southwestern United States: Techniques, Long-term Data Sets, and Trends

Allen et al. 1998

Southwest USA

Multiple

1899–1977

?

Juniper expansion can be observed from repeat photos

https://www.researchgate.net/profile/Thomas_Sisk/publication/235079223_Perspectives_on_the_Land_Use_History_of_North_America_A_Context_for_Understanding_Our_Changing_Environment/links/5886bd5daca272b7b44cd66b/Perspectives-on-the-Land-Use-History-of-North-America

Long Term Vegetation Dynamics of Ponderosa Pine Forests

Bakker 2005 (Dissertation)

Northern AZ

Ponderosa Pine

1941–2004

Grazing on some plots

Overstory cover increased while understory decreased.

https://www.mendeley.com/viewer/?fileId=1326931b-fd88-5527-15f3-a7354491dd5b&documentId=9aa57f69-b3a1-3aa5-9730-a8087b28cfda

Periodic Remeasurement of the Gus Pearson Natural Area

Fule et al., 2001 (Report)

Gus Pearson, AZ

Ponderosa pine

1920–2000

No harvesting

General trend toward declining growth. Increased mortality since 1945.

https://openknowledge.nau.edu/2539/

Comparing Tree-Ring Chronologies and Repeated Timber Inventories as Forest Monitoring Tools

Biondi 1999

Gus Pearson, AZ

Ponderosa pine

1920–1990

No harvesting

Decreased growth rates and higher stand density

https://esajournals.onlinelibrary.wiley.com/doi/abs/10.1890/1051-0761(1999)009%5B0216:CTRCAR%5D2.0.CO%3B2

Twentieth-century changes in forests of the Sierra Nevada

Bouldin 1999

Northern Sierra Nevada, CA

Conifer forests

1935–1992

Fire, drought mortality

Used a large number of forestry plots to show that density of small trees has increased greatly, with decreases in large tree density. Standing dead tree densities are higher than before.

https://www.proquest.com/openview/aac2fa595d16ac697787c3d81b2d330f/1?pq-origsite=gscholar&cbl=18750&diss=y

Stand dynamics and topographic setting influence changes in live tree biomass over a 34-year permanent plot record in a subalpine forest in the Colorado Front Range

Chai et al. 2019

CO

Subalpine forest

1982–2016

insects and pathogens

Despite increased mortality rates over time, there was also a trend of increasing biomass in live trees.

https://doi.org/10.1139/cjfr-2019-0023

Impacts of fire exclusion and recent managed fire on forest structure in old growth Sierra Nevada mixed-conifer forests

Collins et al. 2011

Sierra Nevada, CA

Conifer trees, forb and shrub cover

1911–2007

Fire on some plots

Re-sampled a 1911 timber inventory. Areas with no fire or low severity fire had higher tree density and canopy cover relative to 1911.

esajournals.onlinelibrary.wiley.com/doi/full/10.1890/ES11-00026.1%4010.1002/%28ISSN%292150-8925%28CAT%29VirtualIssue%28VI%29ECS2

Changing forest structure across the landscape of the Sierra Nevada, CA, USA, since the 1930s

Dolanc et al. 2014

Central-Northern Sierra Nevada, CA

Conifer forests

1930s–2000s

Fire in some areas

Used plot data from VTM and FIA to look at changes in forest structure over time (used averages w/in similar regions, not remeasurements of identical plots). Found similar results as Bouldin (above)

https://esajournals.onlinelibrary.wiley.com/doi/full/10.1890/ES14-00103.1

Climate, environment, and disturbance history govern resilience of western North American Forests

Hessburg et al. 2019

Western USA

Multiple

1925–2008 (and other time ranges)

multiple

Review of forest structure change across western USA, including repeat photography.

https://kopernio.com/viewer?doi=10.3389/fevo.2019.00239&route=6

Spatiotemporal Variability of Wildland Fuels in US Northern Rocky Mountain Forests

Keane 2016

Rocky Mountains (MT, ID)

All types of fuels

1993–2004

no disturbance

There is high spatiotemporal variability in fuel accumulation, especially for larger fuels.

http://www.mdpi.com/1999-4907/7/7/129

Long-term surface fuel accumulation in burned and unburned mixed-conifer forests of the Central and Southern Sierra Nevada

Keifer et al. 2006

Sierra Nevada, CA

Fuel loads

1971–2003

Fire on some plots

Fuel load increased over time in unburned plots. In burned plots, 31 years post-fire fuel loads were higher than pre-fire.

http://fireecologyjournal.org/journal/abstract/?abstract=11

A Half Century of Change in Alpine Treeline Patterns at Glacier National Park, Montana, U.S.A.

Klasner and Fagre 2002

MT

High-altitude forests (near treeline)

1945–1991 (some photos from 1927 to 1997)

Road/trail construction/maintenance

Forest homogeneity increased due to greater area with trees and greater tree density within existing patches.

https://doi.org/10.1080/15230430.2002.12003468

Change in Vegetation Patterns Over a Large Forested Landscape Based on Historical and Contemporary Aerial Photography

Lydersen and Collins 2018

Sierra Nevada, CA

Conifer Forest (via aerial photos)

1941–2005

Fire in some areas

The amount of area with dense forest cover increased, and continuous patches of dense forest grew larger.

http://link.springer.com/10.1007/s10021-018-0225-5

Quantifying spatial patterns of tree groups and gaps in mixed-conifer forests: Reference conditions and long-term changes following fire suppression and logging

Lydersen et al. 2013

Sierra Nevada, CA

Conifer trees

1929–2008

Logging, all plots

1929 (pre-logging) was treated as a reference condition, with 2007/8 showing regrowth ~ 80 years after logging. Canopy cover was higher in 2007/8 compared to 1929.

www.sciencedirect.com/science/article/pii/S0378112713003228

Forest vegetation change and surface hydrology following 47 years of managed wildfire

Stevens et al., 2020a

Sierra Nevada, CA

Conifer trees, shrubs

1970–2017

Fire on some plots

Slight increase in total tree density; decrease in density of large trees. Increase in the number of plots with shrub presence.

 https://doi.org/10.1007/s10021-020-00489-5

Southern Cascades Bioregion. Ch. 12 of “Fire in California’s Ecosystems”

Skinner and Taylor, 2018

So. Cascades, CA

Conifer forests

1925–1993

None

Shows increasing tree density, high levels of recruitment, and some mortality of large trees, as well as increased litter cover, using repeat photography.

https://www.degruyter.com/view/books/9780520961913/9780520961913-015/9780520961913-015.xml

Fire regimes and forest changes in mid and upper montane forests of the southern Cascades, Lassen Volcanic National Park, California, U.S.A.

Taylor 2000

So. Cascades, CA

Conifer forests

1925–1993

None

Repeat photography and tree ring records show increased density in Jeffrey Pine and White Fir forests, but little change in red fir forests. Photos also show increased litter cover.

https://doi.org/10.1046/j.1365-2699.2000.00353.x

Long-term response of old-growth stands to varying levels of partial cutting in the Eastside Pine Type

Dolph et al. 1995

East side of Cascades, CA

Conifer forests

1938–1991

None

Repeat measurements show increased density of small trees, but 6% reduction in “sawtimber component”, leading to an overall 13-32% increase in volume of trees over 3.6” DBH.

https://doi.org/10.1093/wjaf/10.3.101

Twenty-year change in aspen dominance in pure aspen and mixed aspen/conifer stands on the Uncompahgre Plateau, Colorado, USA

Smith and Smith 2005

CO

Aspen and conifers

1979–2003

None?

Aspen density remained stable or decreased, while conifer basal area increased over the 20 year study period.

https://doi.org/10.1016/j.foreco.2005.03.018

Vegetation Change and Park Purposes in the High Elevations of Yosemite National Park, California

Vale 1987

Yosemite, CA

High-altitude forests (near treeline)

1900–1985

 

Forests at the upper forest line have increased in density, meadows have been encroached by trees

https://onlinelibrary.wiley.com/doi/abs/10.1111/j.1467-8306.1987.tb00141.x

Widespread Increase of Tree Mortality Rates in the Western United States

van Mantgem et al. 2009

Western USA (CA, OR, WA, ID, CO, AZ)

Forests > 200 years old

~ 1981–2004 (first measurement dates ranged from 1955 to 1994)

Undisturbed

Mortality increased across size classes and elevation ranges. Mean tree density and basal area in the study plots declined slightly during the study period.

https://science.sciencemag.org/content/323/5913/521

Comparison of Historical and Contemporary Forest Structure and Composition on Permanent Plots in Southwestern Ponderosa Pine forests

Moore et al., 2004

AZ and NM

Ponderosa pine

1909–1999

Logging, all plots

Stand density increased greatly; tree diameters shifted toward smaller size classes.

https://academic.oup.com/forestscience/article/50/2/162/4617546

Drought-induced shift of a forest–woodland ecotone: Rapid landscape response to climate variation

Allen and Breshears 1998

New Mexico

Ponderosa pine

1935–1975

drought

ponderosa pine forest receded quickly during a drought

https://www.pnas.org/content/95/25/14839.short

Spatial Patterns of Pinyon–Juniper Woodland Expansion in Central Nevada

Weisberg et al. 2007

Central NV

Pinyon-juniper woodlands

1966–1995

?

Analysis of aerial photos showed an 11% increase in woodland area.

https://doi.org/10.2111/05-224R2.1

Post-1900 Mule Deer Irruptions in the Intermountain West: Principle Causes and Influences

Gruell 1986

Inter-mountain West (ID, MT, NV, UT, WY)

Various woody plants

1868–1982

Grazing and Fire Suppression

Widespread succession from grass dominance to trees and shrubs due to grazing (Reducing competing grass cover) and fire suppression.

https://www.fs.fed.us/rm/pubs_int/int_gtr206.pdf

Vegetation differences in desert shrublands of western Utah’s Pine Valley between 1933 and 1989

Yorks et al. 1992

Western UT

Desert shrubland

1933–1989

Grazing

Canopy cover of grasses greatly increased, with some increase in shrub cover. Density of shrubs decreased (shrubs/m2)

https://journals.uair.arizona.edu/index.php/jrm/article/view/8785

Changes in pinyon-juniper woodlands in western Utah’s Pine Valley between 1933-1989

Yorks et al. 1994

Western UT

Pinyon-juniper woodlands

1933–1989

Grazing

Tree % cover decreased while density increased, due to shift in tree cover from juniper to pinyon (narrower crown). Shrub and grass % cover increased. Shrub density increased.

https://repository.arizona.edu/bitstream/handle/10150/644369/8955-8836-1-PB.pdf?sequence=1

A century of vegetation change in the San Juan Mountains, Colorado: An analysis using repeat photography

Zier and Baker 2006

CO

Conifer and aspen forests, grasslands

~1900–~2000

Varying disturbances

Both conifers and deciduous trees increased in extent, partially as recovery from disturbances. There was some encroachment of trees into grass/shrublands.

https://doi.org/10.1016/j.foreco.2006.02.049

Fire and vegetative trends in the Northern Rockies: interpretations from 1871-1982 photographs

Gruell 1983

Northern Rockies, MT

Forest

1871–1982

 

Woody vegetation increased due to suppressed wildfire

https://www.fs.usda.gov/treesearch/pubs/32994

ENVIRONMENTAL CHANGE IN GLACIER NATIONAL PARK, MONTANA: AN ASSESSMENT THROUGH REPEAT PHOTOGRAPHY FROM FIRE LOOKOUTS

Butler and DeChano 2001

MT

Montane forest

1935–1990s

Avalanches, glacial recession, anthropogenic developments

Increased forest density and extent likely due to a combination of fire suppression and climate change (including receding glaciers)

https://doi.org/10.1080/02723646.2001.10642744

Fire’s Influence on Wildlife Habitat on the Bridger-Teton National Forest, Wyoming - Volume I: Photographic Record and Analysis

Gruell 1980

WY

Multiple tree and shrub types

1872–1975

Some fires, mostly prior to 1941

Increases in conifers and sagebrush

https://digitalcommons.usu.edu/barkbeetles/95/

100,000 Trees Can’t Be Wrong: Permanent Study Plots and the Value of Time

Duncan 2004 (FS publication for managers)

Pacific NW (OR, WA)

Multiple forest types

1910–2002

Varying disturbances

There is a network of long term forestry plots in OR and WA, managed by the PNW region of the USFS. Biomass accumulation can continue even after 80 years post-disturbance.

https://www.fs.usda.gov/treesearch/pubs/6956

Thirty-six years of tree population change in an old-growth Pseudotsuga- Tsuga forest

Franklin and DeBell 1988

Cascade Range, WA

Conifers

1947–1983

Undisturbed

Old growth forest showed slight decline in tree density. The diameter distribution shifted upward, and 22% of the original stems died during the study period (though this was almost matched by recruitment)

https://doi.org/10.1139/x88-093

Carbon stocks and accumulation rates in Pacific Northwest forests: role of stand age, plant community, and productivity

Gray et al., 2016

Pacific NW (OR, WA, CA, ID)

Live and dead trees, many types

1993–2007

disturbance on some plots

Older trees accumulate C slower, but forests still have net C increase until ~ 400 years old when high mortality outweighs growth

http://doi.wiley.com/10.1002/ecs2.1224

Input and decay of coarse woody debris in coniferous stands in western Oregon and Washington

Sollins 1982

Pacific NW (OR,WA)

Coarse woody debris

16–46 year span

Undisturbed

Tree mortality resulted in dry-matter transfer of 1.5–4.5 Mg•ha − 1•year− 1 of boles and branches to the forest floor and 0.3–1.3 Mg•ha − 1•year− 1 of large-diameter roots directly to the mineral soil. , with decay rates slower than accumulation.

http://www.nrcresearchpress.com/doi/10.1139/x82-003

Historical and current landscape-scale ponderosa pine and mixed conifer forest structure in the Southern Sierra Nevada

Stephens et al. 2015

Southern Sierra Nevadas, CA

Ponderosa pine and mixed conifer forest

1911–2005

Very little disturbance

Compared tree density and canopy cover measured by 1911 surveys and modern FIA inventories in same area, though not the same exact plot locations. Found increases in tree density, fir dominance, and canopy cover.

https://esajournals.onlinelibrary.wiley.com/doi/full/10.1890/ES14-00379.1

Vegetation and Soil Condition Changes on a Subalpine Grassland in Eastern Oregon

Strickler 1961

Wallowa Mountains, OR

Grasslands/shrubs

1938–1956

Grazing

Total biomass and veg. cover increased over the 20 years due to improved range management.

https://www.fs.fed.us/pnw/pubs/pnw_os_rp-40.pdf